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The gdh2 / gdh2 mutant failed to utilise arginine and proline as both the sole carbon and nitrogen sources

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Shi F, Zhang Z, Cui H, Wang J, Wang Y, Tang Y, et al

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Deficiency Symptoms Men and women fed intravenously (IV) with solutions that contained adequate methionine and folate but lacked choline have been found to develop a condition called nonalcoholic fatty liver disease (NAFLD) and signs of liver damage that resolved when choline was provided (12)